AQs are over all too strong and with them slowly more and more in game, the normal queens are less and less good in comparison. It is very hard to gather team for common queen matching both landform of all 12 bees and either like or dislike of queen, which is part of top tier end-game to maximize team power. AQs remove from equation whole one part, the landform, as they provide full bonus to all bees no matter what is their specialty. This basically killed normal queen market. To balance the this issue a bit, I would suggest this change to AQ mechanics:
Ancient Queens newly provide their landform siege bonuses ONLY to bees that match either like or dislike of the queen, not-matching bees receive bonus as if the AQ was normal queen. This will motivate players to gather like or dislike matching bees and will improve marketplace activity and will reduce overall OP of AQs.